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Updated on
27 September

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Question about English (US)

What is the subject of 'running' in this sentence? 'The motels' or 'north-south routes?'

"The motels clustered along transcontinental highways such as U.S. Routes 40 and 66 and along the north-south routes running up and down both the east and west coasts."
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What is the subject of 'running' in this sentence? 'The motels' or 'north-south routes?'
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